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Classification
Urytalpa Edwards, 1929
Distribution of Urytalpa vicina
Distribution of Urytalpa rhapsodica
Distribution of Urytalpa nussbaumi
Urytalpa trivittata (Lundström, 1914)
General description:Description based on Kjærandsen et al. (2009): On the genus Urytalpa Edwards (Diptera: Keroplatidae) in the Nordic and Nearctic regions, with fixation of a new type species and a key to world males
Diagnostic description:Most similar to U. dorsalis but differs by having more slender and less petiolated and less laterally compressed abdomen and by having the thoracic stripes distinctly separated. The larger apicomedial outgrowth of the gonocoxite with 5–6 strong setae in a row is a distinctive character for this species. Urytalpa dorsalis and U. trivittata differ from the other Nordic species in having gonocoxites without larger outgrowths and the gonostylus is retracted dorsally into the gonocoxites. In U. trivittata, however, the gonostylus is subtriangular in lateral view and somewhat larger. The female can be separated from U. dorsalis and U. macrocera by the combination of an oblong and petiolated cercus, sternite VIII having truncated apicomedial corners with a narrow setose excavation, and by having distinctly ovate spermathecae with a relatively thin inner wall.
Morphology:Male
Coloration (specimen in alcohol). Head brown with mouth parts paler. Antenna brown, scape, pedicel and basal part of first flagellomere paler. Thorax yellow with pale antepronotum; dark brown mesonotal stripes distinctly separated,narrowly surrounded by pale yellow ground; humeral area pale, extending laterally to behind wing base. Wings clear, yellow tinged, veins brown. Halter pale. Legs slightly paler than thorax. Abdomen brown;tergites I–V with narrow pale bands apically; tergites VI–IX dark brown; sternites I–V pale; terminalia yellowish brown.
Terminalia (Fig.). Gonocoxite connected ventrally by a narrow strip only; apically with a distinct apicomedial outgrowth with 5–6 strong setae in a row (A). Large hypandrial lobe forming thin hyaline plate ventromedially (A). Gonostylus retracted into gonocoxite, small and subtriangular in lateral view with the tip pointing dorsally (D). Aedeagal apparatus long but not high, with less compressed plate extending anteriorly into segment V, and long associated apodemes laterally. Tergite IX without outgrowths, apical corner pointed ventrally, without patch of short internal setae; with shallow posterior and deep anterior U-shaped incision (B). Proctiger small and short, cercus broadly ovate in lateral view (A).
Female.
Coloration. As in male with dark brown mesonotal stripes distinctly separated.
Terminalia (Fig.). Cercus oblong and petiolated (A.). Sternite VIII with truncated corners, with a narrow excavation apicomedially covered with short stiff setae. Spermathecae distinctly ovate, spermathecae and ducts with medium thick inner walls (B.).
Size:Male (n=5). Body length 6.5–8.5, 7.1 (n=10) mm. Wing length 5.14–5.88, 5.5 mm, or3.24–3.64, 3.39 as long as profemur. Antenna length 2.2–2.64 (n=4) mm.
Female (n=1). Body length 7.5 mm. Wing length 6 mm. Antenna length 1.82–1.86 mm.
Distribution:European; with scattered records only from Finland, Germany, Italy, Norway and Sweden.Possibly boreal-mountainous with main distribution in northern parts of the boreal zone.
Urytalpa macrocera (Edwards, 1913)
General description:Description based on Kjærandsen et al. (2009): On the genus Urytalpa Edwards (Diptera: Keroplatidae) in the Nordic and Nearctic regions, with fixation of a new type species and a key to world males
Diagnostic description:The long slender antenna, uniform dark brown coloration, and rounder wing with shorter R-Mvein fusion easily distinguishes this species from the other members of the genus. Thelong slender apical projection of the gonocoxites is shared only with U. galdes, sp. n. but mostother structures of the male terminalia are unique and distinct, including the large spoon shaped projectionlaterally from tergite IX with a dense brush of long setae internally. The female can be separatedfrom U. trivittata and U. dorsalis by the short, truncated abdomen and by having globularspermathecae with thin inner walls and thin ducts.
Morphology:Male
Coloration (specimen in alcohol). Head dark brown, mouth parts brown. Antenna brown with basal part of first flagellomere pale.Thorax dark brown; thoracic stripes dark brown, diffusely separated; humeral area brown. Wings clear, yellowtinged, veins paler brown than in other species. Halter whitish. Legs brown. Abdomen uniformly dark brown.
Terminalia (Fig.). Gonocoxites divided ventrally, with long slender projection. Hypandrial lobe tongue shaped and curved dorsally. Double "bladder shaped" aedeagal guides situated above the gonocoxite. Gonostylus long and slender, subrectangular in lateral view with the apicodorsal corner slightly elongated. Aedeagal apparatus short and small compared to other species, with compressed plate extending only slightly beyond apex of the stout and short lateral apodeme. Tergite IX with large spoon shaped projection laterally and with large dense brush of long setae internally, with deep posterior and shallow anterior U-shaped incision. Proctiger small and short, cercus sessile, broadly ovate in lateral view.
Female
Coloration. Overall brown as in male.
Terminalia (Fig.).Tergites VIII and IX weakly sclerotized apically in front of cercus. Cercus sessile and ovate (A.). Sternite VIII divided ventrally, with slightly truncated corner, with a narrow excavation apicomedially covered with short stiff setae (B.). Spermathecae globular, spermathecae and ducts with thin inner walls (A.).
Size:Male (n=5). Body length 6.5–8.5, 7.79 (n=7) mm. Wing length 5–6.72. 5.83 mm, or3.74–4.15, 3.89 as long as profemur. Antenna length 2.84–3.9 (n=4) mm.
Female (n=3). Body length 5–5.5 mm. Wing length 4.04–4.08 mm, or 3.29–3.54 as long as profemur. Antenna length 1.62–1.7 mm.
Distribution:Western Europe; known only from northern Britain (Scotland and northern England),France, Norway, Sweden and The Netherlands.
Urytalpa dorsalis (Staeger, 1840)
General description:Description based on Kjærandsen et al (2009): On the genus Urytalpa Edwards (Diptera: Keroplatidae) in the Nordic and Nearctic regions, with fixation of a new type species and a key to world males
Diagnostic description:This species comes closest to U. trivittata and U. atriceps in shape of the terminalia, but differs by having the thoracic stripes darker and fused in males. The gonostylus is small, tapered and claw-like in lateralview, retracted and directed dorsad to the extent that it is almost totally hidden in ventral view. Such a small, retracted gonostylus is also found in U. trivittata and U. sapporoensis (Okada, 1938).This is the only Nordic species of Urytalpa where the male abdomen due to the extra large anterior extension of the aedeagal apparatus appears distinctly petiolated and laterally compressed. The female can be separated from U. trivittata and U. macrocera by the combination of a short ovate sessile cercus, sternite VIII having convex apicomedial corners with a larger setose excavation, and by having globular spermathecae with thick inner wall.
Morphology:Male (n=5). Body length 6–9, 7.5 (n=7) mm. Wing length 4.46–6.02, 5.24 mm, or3.28–3.44, 3.36 as long as profemur. Antenna length 2.18–2.48, 2.32 mm.
Coloration (specimen in alcohol). Head dark brown, mouthparts and palps pale yellowish. Antenna dark brown, scape, pedicel and basal part of first flagellomere pale. Thorax brown with antepronotum pale; preepisternum 2 with darker lower half; laterotergite with darker posteroventral part; mesonotal stripes dark, almost completely fused,separated anteriorly; humeral area pale, extending laterally to behind wing base. Wings clear, yellow tinged,veins brown. Halter pale. Legs pale whitish yellow. Abdomen dark brown with variable amount of yellow;tergites I–V usually with large triangular apicolateral yellow marking; tergites VI–IX darker, with at most small yellow patches laterally on tergites VI–VII; sternites I–VI pale; terminalia dark brown.
Terminalia. (Fig) Gonocoxites fused ventrally by a broad connection; apically with a short apicomedial outgrowth without strong setae. Large hypandrial lobe forming thin hyaline plate ventromedially. Gonostylus retracted into gonocoxite, small and claw like in lateral view with the tip pointing apicoventrally. Aedeagal apparatusvery large, with compressed plate large and tall, extending anteriorly into segment V, and long associated apodemes laterally. Tergite IX without outgrowths, apical corner with patch of short internal setae; with shallow posterior and deep anterior U-shaped incision. Proctiger small and short, cercus oblong in lateralview.
Female (n=2). Body length 7.5 mm. Wing length 5.68–5.9 mm, or 3.38–3.6 as long as profemur. Antennalength 1.75 mm.
Coloration. Thorax usually all yellow without dark markings; abdomen variablefrom all dark to entirely yellow; otherwise as in male.
Terminalia. Cercus short ovate andsessile. Sternite VIII with convex corners, with a large excavation apicomedially covered with short stiff setae. Spermathecae globular, spermathecae and ducts with thick inner walls.
Distribution:Widespread in Europe; recorded from Belgium, Bulgaria, Czech Republic, Denmark,Finland, France, Germany, Great Britain, Hungary, Ireland, Italy, Latvia, Netherlands, Norway, Poland,Romania, Russia (NET), Slovakia, Spain, Sweden and Switzerland.
Urytalpa atriceps (Edwards,1913)
General description:Description based on Kjærandsen et al (2009): On the genus Urytalpa Edwards (Diptera: Keroplatidae) in the Nordic and Nearctic regions, with fixation of a new type species and a key to world males
Diagnostic description:This species shares terminalia with fused gonocoxites and very long aedeagal apparatus with U. dorsalis and U. trivittata. It can be distinguished from these and other species by having long, digitateprojections from tergite IX and relatively large and exposed gonostyli. The shape of the bifidgonostylus is unique with a brush of strong setae on the inner smaller branch
Morphology:Male
Coloration (dry, pinned specimen). Head dark blackish brown with pale yellowishbrown mouth parts and palps. Antenna dark brown; scape, pedicel and basal part of first flagellomere paleryellow. Thorax yellow with mediotergite dark brown; preepisternum with brown posterior part and anterioredge; thoracic stripes brown, narrowly surrounded by pale yellow ground, humeral area pale, extending laterally to behind wing base. Wings clear, yellow tinged. Halter yellow. Legs yellow. Abdomen yellow withtergites VI–IX dark brown, sternites VII–VIII brownish yellow.
Terminalia (Fig.). Gonocoxites connectedby a narrow strip and prolonged apicolaterally (A). Gonostylus bifurcate, with a broad base; ventral branch strong,ending in a sclerotized fold; dorsal branch shorter with about 5 long strong setae forming a brush apically (D) . Aedeagal apparatus long but not high, with compressed plate extending anteriorly into segment V, and long associated apodemes laterally. Tergite IX with a long, narrow, digitate projection apicolaterally without patchof short internal setae; with deep posterior and anterior U-shaped incision (B&C). Proctiger small and short, cercus thin, rhombic in lateral view (B&C).
Female unknown.
Size:Male (n=1). Body length 6 mm. Wing length 4.84 mm or 3.36 as long as profemur. Antenna length 2.26 mm.
Female. Unknown
Distribution:North-western Europe; known only from southern England, the Netherlands, Norway and Sweden.
Distribution of Urytalpa juliae
Distribution of Urytalpa galdes
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