Tarnania dziedzickii
In caves
In caves
in caves
The text is based on Kjærandsen (2006): Review of fungus gnats in the genus Tarnania Tuomikoksi, with a phylogeny of the Rymosia s.l. genus group (Diptera : Mycetophilidae)
Apart from the coloration as given in the key (• males) the male can be separated from other Tarnania primarily by the shape of the hypandrial lobe, which in ventral view is subrectangular and parallel-sided for most of its length, except at extreme tip where it is constricted to a tiny, pointed apex (A). Further diagnostic characters of the male include: The ventral branch of the gonostylus usually having a distinct protrusion apicodorsally and short setae internally on apical half (A); the median branch being evenly broad towards apex (A); the cercus being slender, without dorsal widening basomedially (A); and the aedeagus being narrow triangular (A). The female is best separated from other Tarnania by the combination of dark coloration (females), having narrowly pointed gonapophysis VIII and long, slender cercus II (A).
Male (n = 5, except where otherwise stated):
Coloration (fresh animal in alcohol): Antenna with scape and pedicel light brown, basal half of first flagellomere light yellow, rest of flagellum uniformly brown. Head reddish brown, also on postgena; clypeus reddish brown, maxillary palp yellow. Thorax with sharply defined mesonotal stripes narrowly surrounded by pale yellow ground graduated towards darker reddish brown humeral areas; Antepronotum, proepisternum, anepisternum, lower two thirds of katepisternum, upper part of anepimeron, laterotergite, mediotergite and metepisternum reddish brown. Wings unmarked; yellow tinted, especially in costal and radial sector; wing veins yellow with dark setae. Halter whitish. Legs mainly yellow; fore coxa brownish basally; mid and hind coxae yellow, with large but diffuse brownish spot medially. Abdomen mainly reddish brown with narrow yellow bands apically, tergites III and IV with reddish yellow triangular area lateroapically granulated with large dark spots; tergite VI dark brown; sternites gradually darker towards terminalia; terminalia yellow basally, reddish brown apically; tergite IX and cercus yellow.
Head: Round, width/length to frontal tubercle 1.26–1.54, 1.41. Antenna 1.5–1.86 (n=3) mm long. First flagellomere 1.62–1.81, 1.72 times as long as second flagellomere. Second flagellomere 1.5–1.7, 1.6 times as long as wide. Length/width of clypeus 0.66–0.89, 0.76. Antepenultimate palpomere 0.15–0.19, 0.17 mm long, palpomere ratios 1: 1.23–1.41, 1.29: 2.73–3.04, 2.88.
Thorax: One strong proepisternal bristle.
Wings: Wing length to length of R1 2–2.17, 2.09. R5 slightly sinusoid, wing length to length of R5 1.61–1.68, 1.65. Length of ta to length of M-petiole 1.15–1.43, 1.29. Fork length ratio (A/B) 0.99–1.09, 1.05. Fork width ratios (C/D and E/F) 2–2.4, 2.17 and 0.7–0.86, 0.76. M-ratios 0.58–0.64, 0.61 and 0.64–0.7, 0.66. CuA-ratios 0.69–0.84, 0.75 and 0.9–1.09, 0.97. CuP length to length of wing 0.55–0.69, 0.59. A1 length to length of wing 0.45–0.48, 0.47.
Legs: Leg ratios given for fore, mid and hind leg: LR 0.95–1.03, 0.99: 0.97–1.04, 0.99: 0.75–0.79, 0.78; SV 1.94–2.1, 2.01: 1.92–2.04, 1.99: 2.29–2.39, 2.34; BV 1.64–1.81, 1.76: 2.02–2.15, 2.07: 2.66–2.91, 2.76; TR 1.48–1.63, 1.55: 1.74–1.91, 1.82: 2.11–2.25, 2.16.
Terminalia (•): Genitalia as large as or larger than abdominal segment VI. Tergite IX distinctly shorter than wide, covered with relatively few setae. Cercus narrow, slender, 5–5.94 (n=4) as long as wide in dorsal view; without dorsal widening basomedially. Aedeagus narrow triangular, with deep medial invagination apically. Hypandrial lobe in ventral view a long subrectangular and parallel-sided plate, without constriction except at extreme tip; in lateral view with small, distinct hook apically. Gonostylus with dorsointernal branch fan-shaped; median branch evenly broad towards apex, apically with short, lanceolate seta; ventral branch usually with distinct protrusion apicodorsally, with short setae internally on apical half.
Female (n = 5, except where otherwise stated):
Coloration (fresh animal in alcohol): As for male, except abdominal tergites III - VI with larger, subrectangular reddish yellow areas laterally, more heavily granulated with large dark spots; tergite VII and terminalia reddish yellow.
Head: Round, width/length to frontal tubercle 1.22–1.56, 1.39. Antenna short, 1.8–2.2, 1.96 mm long. First flagellomere 1.62–2.09, 1.82 times as long as second flagellomere. Second flagellomere 1.6–1.77, 1.68 times as long as wide. Length/width of clypeus 0.64–0.77, 0.7. Antepenultimate palpomere 0.16–0.21, 0.19 mm long, palpomere ratios 1: 1.23–1.5, 1.37: 2.58–3.1, 2.83.
Thorax: One strong proepisternal bristle.
Wings: Wing length to length of R1 1.98–2.09, 2.04. R5 slightly sinusoid, wing length to length of R5 1.61–1.67, 1.63. Crossvein ta with small, distinct white spot. Length of ta to length of M-petiole 1.05–1.43, 1.27. Fork length ratio (A/B) 0.82–1.14, 1.02. Fork width ratios (C/D and E/F) 1.79–2.33, 2.15 and 0.57–0.79, 0.68. M-ratios 0.57–0.59, 0.58 and 0.62–0.65, 0.63. CuA-ratios 0.63–1.01, 0.75 and 0.8–1.33, 0.98. CuP length to length of wing 0.55–0.58, 0.57. A1 length to length of wing 0.43–0.46, 0.45.
Legs: Leg ratios given for fore, mid and hind leg: LR 0.97–1.02, 0.99: 0.98–1.01, 0.99: 0.77–0.85, 0.79; SV 1.98–2.03, 2.01: 1.97–2.05, 2: 2.22–2.4, 2.31; BV 1.72–1.91, 1.82: 2.01–2.14, 2.1: 2.75–2.98, 2.88; TR 1.56–1.71, 1.62: 1.74–1.88, 1.82: 2.19–2.31, 2.24.
Terminalia (•): Cercus I conical. Cercus II slender, 3.53–4.58, 4.29 as long as wide in lateral view. Gonapophysis VIII narrowly pointed towards apex; apicodorsal margin usually distinct sinusoid.
Male (n = 5): Total length 7.00 – 8.00, 7.30 mm. Wing length 3.84 – 4.38, 4.12 mm, or 3.33 – 3.47, 3.4 x as long as profemur. Mesonotum length 1.04 – 1.24, 1.11 mm, or 0.26 – 0.28, 0.27 x as long as wing.
Female (n = 5): Total length 7.50 – 9.50, 8.20 mm. Wing length 4.20 – 5.24, 4.71 mm, or 3.23 – 3.53, 3.41 x as long as profemur. Mesonotum length 1.14 – 1.50, 1.32 mm, or 0.27 – 0.29, 0.28 x as long as wing.
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