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Classification
Tarnania tarnanii (Dziedzicki, 1910)
Tarnania tarnanii
Habitat:In caves
Tarnania tarnanii
Distribution:After Polevoi & Barkalov, 2017
Tarnania tarnanii (Dziedzicki, 1910)
General description:The text is based on Kjærandsen (2006): Review of fungus gnats in the genus Tarnania Tuomikoksi, with a phylogeny of the Rymosia s.l. genus group (Diptera : Mycetophilidae)
Diagnostic description:Apart from the coloration as given in the key (• males) the male can be separated from other Tarnania primarily by the shape of the hypandrial lobe, which in ventral view is wide rectangular basally, then abruptly constricted into a short, pointed plate (D); in lateral view with less distinct apical hook (D). Further diagnostic characters of the male include: One strong and one lesser propleural bristles; ventral branch of the gonostylus internally with a cushion, densely covered with long setae (D); dorsointernal branch being more spatulate than fan-shaped (D); and the aedeagus being subquadrate (D). The female is best separated from other Tarnania by the combination of pale coloration (females), having narrowly pointed gonapophysis VIII, and short, ovate cercus II (D).
Morphology:Male (n = 6, except where otherwise stated):
Coloration (fresh animal in alcohol): Antenna with scape, pedicel and basal half of first flagellomere light yellow; rest of flagellum reddish yellow, darker towards apex. Head reddish yellow, darker in front of lateral ocellus; postgena pale yellow; clypeus yellow, maxillary palp yellow. Thorax reddish yellow without well defined mesonotal stripes, humeral areas pale yellow; most of anepisternum, upper part of anepimeron, lower half of katepisternum, posterior margin of laterotergite, mediotergite and metepisternum yellowish brown; thoracic sclerites otherwise pale yellow. Wing unmarked; yellow tinted, especially in costal and radial sector; wing veins yellow with dark setae. Halter whitish. Legs pale yellow; mid and hind coxae with medium sized, distinct brownish spot apicomedially. Abdomen mainly reddish brown with narrow yellow bands apically, tergites II - IV with triangular yellow area lateroapically, hardly granulated with dark spots except in narrow transition zone; tergite VI brown; sternites gradually darker towards terminalia; terminalia yellow; tergite IX and cercus yellow.
Head: Round, width/length to frontal tubercle 1.31–1.41, 1.37 (n=5). Antenna 1.4–1.52, 1.47 (n=5) mm long. First flagellomere 1.79–3.44, 2.32 (n=5) times as long as second flagellomere. Second flagellomere 0.73–1.19, 1.03 (n=5) times as long as wide. Length/width of clypeus 0.66 – 0.87, 0.77 (n=5). Antepenultimate palpomere 0.15 – 0.17, 0.16 (n=5) mm long, palpomere ratios 1: 1.33–1.64, 1.41 (n=5): 2.84–3.56, 3.15 (n=5).
Thorax: One strong and one less strong propleural bristles.
Wings: Wing length to length of R1 1.98–2.26, 2.16. R5 slightly sinusoid, wing length to length of R5 1.68–1.74, 1.71. Crossvein ta with small, distinct white spot. Length of ta to length of M-petiole 1.25–1.5, 1.35. Fork length ratio (A/B) 1.1 – 1.21, 1.17. Fork width ratios (C/D and E/F) 1.55 – 1.84, 1.75 and 0.69–0.92, 0.81. M-ratios 0.61 – 0.63, 0.62 and 0.67–0.69, 0.68. CuA-ratios 0.61 – 0.71, 0.65 and 0.79–0.93, 0.84. CuP length to length of wing 0.55–0.6, 0.57. A1 length to length of wing 0.44–0.49, 0.46.
Legs: Leg ratios given for fore, mid and hind leg: LR 1.02–1.07, 1.05 (n=5): 1.04–1.11, 1.06 (n=5): 0.72–0.8, 0.76 (n=5); SV 1.83–1.95, 1.89 (n=5): 1.78–1.92, 1.87 (n=5): 2.33–2.45, 2.39 (n=5); BV 1.32–1.38, 1.35 (n=5): 1.65–1.88, 1.79 (n=5): 2.57–2.89, 2.75 (n=5); TR 1.28–1.4, 1.33 (n=5): 1.63–1.74, 1.7 (n=5): 1.93–2.19, 2.09 (n=5).
Terminalia (•): Genitalia smaller than abdominal segment VI. Tergite IX subrectangular, shorter than wide, with relatively few setae. Cercus short, 2.86–3.37, 3.16 (n=5) as long as wide in dorsal view; with dorsal widening basomedially. Aedeagus subquadrate. Hypandrial lobe in ventral view wide rectangular basally, then abruptly constricted into a short, pointed plate; in lateral view with less distinct apical hook. Gonostylus with dorsointernal branch more spatulate than fan-shaped; median branch with apical half distinctly tapered, apically with strong seta; ventral branch without distinct protrusion apicodorsally, internally with a cushion, densely covered with long setae.
Female (n = 6, except where otherwise stated):
Coloration (fresh animal in alcohol): As for male, except abdominal tergites II - V with larger, triangular yellow areas laterally; tergite VII and terminalia reddish yellow.
Head: Round, width/length to frontal tubercle 1.29–1.52, 1.4 (n=5). Antenna 1.28–1.7, 1.47 (n=5) mm long. First flagellomere 1.79–3.44, 1.99 (n=5) times as long as second flagellomere. Second flagellomere 1.04–1.3, 1.13 (n=5) times as long as wide. Length/width of clypeus 0.66 – 0.74, 0.69 (n=5). Antepenultimate palpomere 0.15 – 0.2, 0.17 (n=5) mm long, palpomere ratios 1: 1.28–1.44, 1.38 (n=5): 2.55–3.02, 2.86 (n=5).
Thorax: One strong and one lesser strong propleural bristles.
Wings: Wing length to length of R1 2.1–2.26, 2.16. R5 slightly sinusoid, wing length to length of R5 1.62–1.8, 1.69. Crossvein ta with small, distinct white spot. Length of ta to length of M-petiole 1.14–1.7, 1.41. Fork length ratio (A/B) 1.05 – 1.27, 1.15. Fork width ratios (C/D and E/F) 1.6 – 2, 1.83 and 0.65–0.81, 0.71. M-ratios 0.53–0.58, 0.56 and 0.58–0.65, 0.62. CuA-ratios 0.52–0.67, 0.61 and 0.65–0.83, 0.76. CuP length to length of wing 0.55–0.62, 0.59. A1 length to length of wing 0.44–0.47, 0.45.
Legs: Leg ratios given for fore, mid and hind leg: LR 1.02–1.07, 1.04 (n=5): 1.01–1.04, 1.03 (n=5): 0.74–0.78, 0.75 (n=5); SV 1.84–1.97, 1.92 (n=5): 1.86–1.93, 1.9 (n=5): 2.36–2.46, 2.41 (n=5); BV 1.34–1.41, 1.37 (n=5): 1.79–1.85, 1.83 (n=5): 2.55–2.82, 2.72 (n=5); TR 1.32 – 1.44, 1.36 (n=5): 1.64–1.75, 1.67 (n=5): 2 – 2.14, 2.08 (n=5).
Terminalia (•): Cercus I long, less conical. Cercus II short, ovate, 2.31–2.73, 2.57 (n=5) as long as wide in lateral view. Gonapophysis VIII narrowly pointed towards apex; apicodorsal margin less sinusoid.
Size:Male (n = 6): Total length 6–7.5, 6.7 mm. Wing length 3.68–3.94, 3.85 mm, or 3.35–3.61, 3.48 x as long as profemur. Mesonotum length 1.04 – 1.12, 1.08 mm, or 0.27– 0.29, 0.28 x as long as wing.
Female (n = 6): Total length 7–8.5, 7.4 mm. Wing length 3.74 – 4.62, 4.16 mm, or 3.63–3.91, 3.71 x as long as profemur. Mesonotum length 1–1.3, 1.15 mm, or 0.27–0.29, 0.28 x as long as wing.
Tarnania tarnanii, distribution map
